BFS Plants


Status of the list. Please note that this is a “living” list, constantly being refined and updated. Plant species and generic circumscriptions may change as a result of molecular research. In addition, some plant groups at the BFS are particularly challenging and open to revision. We attempt to keep the list up-to-date via the Jepson eFlora, and we follow that nomenclature unless otherwise noted.

Species that have reported at the BFS but whose presence has not been confirmed have been moved to a separate list. If you can confirm that any of these species occur (or have occurred) at the BFS, please notify the webmaster.

Conservation status. According to State and Federally Listed Endangered, Threatened, and Rare Plants of California, April 2021 (California Department of Fish & Wildlife), only one plant species at the BFS has official conservation status:

  • Berberis nevinii (Nevin’s Barberry) – on both State and Federal Endangered Species Lists.

Several other BFS plant species are listed as rare or threatened and are being tracked in the California Native Biodiversity Database, but have no official protection. See Special Vascular Plants, Bryophytes, and Lichens List, April 2021 (California Department of Fish & Wildlife) for details:

  • Horkelia cuneata ssp. puberula (Mesa Horkelia) – Critically imperiled
  • Lepidium virginicum var. robinsonii (Robinson’s Peppergrass) – Vulnerable
  • Deinandra paniculata (Paniculate Tarplant) – Limited distribution
  • Iva hayesiana (San Diego Marsh-Elder) – Imperiled
  • Juglans californica (Southern California Black Walnut) – Limited distribution

Photos & descriptions. ‘BFS’ links will show plants photographed at the BFS. We eventually hope to have photographs of all the plants on the site. Photos on the BFS pages may be used for any personal or academic purposes as long as notification of use is sent to the BFS Webmaster and the image is clearly credited with ©[Name of photographer]. For higher-resolution images, please contact the photographer. Note that no photos taken at the BFS may be used for commercial purposes without the express permission of the BFS Director.

‘CP’ links will return all photographs of the listed taxon in the CalPhotos, Digital Library Project database. Note that you must check the stipulations with each photograph for permissible uses of the CalPhotos images. ‘Jep’ links will return the treatment and bioregional distribution for a taxon from Jepson eFlora.

Origin. The origin listing indicates whether a plant is thought to be indigenous to the site (Native), a California native that is not indigenous but has naturalized or been planted on the BFS (Native*), or not native to California (Non-native).

Plants designated NativeL are riparian or aquatic plants around or in pHake Lake. After lake construction was completed in 1978, BFS faculty, in consultation with Professor Robert Thorne of the Rancho Santa Ana Botanic Garden, introduced a variety of riparian and aquatic native plants. Additional native plants have appeared on their own via birds or aerial seed dispersal.

Validation. For many BFS plants, voucher specimens have been deposited in herbaria, and links are provided to online data for those specimens. If no online herbaria data are available, “photo” indicates that photos of the plant are available via the “BFS” link under “Photos”. If neither voucher specimens nor photos exist, but the identification is from a reliable source, initials indicate the source.

Non-native plants around the old infirmary. The area surrounding the former Claremont Colleges’ infirmary building was once planted with a wide variety of plants, most of which were non-native. Most of these were removed and replaced with native plants during the renovation of the building for the Redford Conservancy; however, those that were identified remain on the list with a note.

London RocketSisymbrium irioCP, Jepnon-nativeSD

Common name Scientific name Photos &
Origin* Validation* Notes


Muskroot Family (Adoxaceae)

Blue Elderberry Sambucus nigra ssp. caerulea BFS, CP, Jep native RSA549568
(above all as Sambucus mexicana)
Formerly Sambucus mexicana. Formerly placed in Caprifoliaceae.

Amaranth Family (Amaranthaceae)

Tumbleweed Amaranthus albus BFS, CP, Jep non-native Photo, SD
Procumbent Pigweed Amaranthus blitoides CP, Jep native RSA766851
Red Amaranth Amaranthus cruentus BFS, CP, Jep non-native Photo Has appeared as a waif.

Sumac or Cashew Family (Anacardiaceae)

Laurel Sumac Malosma laurina BFS, CP, Jep native Photo
Skunk Bush Rhus aromatica BFS, CP, Jep native RSA549571
(above all as Rhus trilobata)
Formerly Rhus trilobata.
Lemonade Berry Rhus integrifolia BFS, CP, Jep native RSA549577
Sugar Bush Rhus ovata BFS, CP, Jep native RSA549578
Peruvian Pepper Tree Schinus molle BFS, CP, Jep non-native RSA611984
Western Poison Oak Toxicodendron diversilobum BFS, CP, Jep native RSA614157

Carrot Family (Apiaceae)

Wild Celery Apiastrum angustifolium BFS, CP, Jep native RSA748786
Rattlesnake Weed Daucus pusillus BFS, CP, Jep native RSA616156

Dogbane Family (Apocynaceae)

Kotolo, Woolypod Milkweed Asclepias eriocarpa BFS, CP, Jep native RSA764402 Formerly placed in Asclepiadaceae.
Climbing Milkweed Funastrum cynanchoides var. hartwegii BFS, CP, Jep native Photo Formerly Sarcostemma cynanchoides ssp. hartwegii. Formerly placed in Asclepiadaceae.
Common Oleander Nerium oleander BFS, CP, Jep non-native Photo One on east side of lake and several along the Foothill and Mills fences.

Sunflower Family (Asteraceae)

Annual Bur-Sage Ambrosia acanthicarpa CP, Jep native JG, SD
California Sagebrush Artemisia californica BFS, CP, Jep native Photo
Mugwort Artemisia douglasiana BFS, CP, Jep native SD
Coyote Brush Baccharis pilularis BFS, CP, Jep native RSA549569
Mule Fat Baccharis salicifolia subsp. salicifolia BFS, CP, Jep native RSA611982
Common Beggar-Ticks Bidens pilosa BFS, CP, Jep non-native RSA823057 A single plant (since removed) was spotted in the following summer in the area burned in the 2013 Foothill Fire.
California Brickellbush Brickellia californica BFS, CP, Jep native Photo On plant by the toad pond perished in the prolonged drought of 2014-2017, but at least one other is present in the ‘Neck’.
Italian Thistle Carduus pycnocephalus subsp. pycnocephalus BFS, CP, Jep non-native RSA751114
Tocalote, Maltese Star-Thistle Centaurea melitensis BFS, CP, Jep non-native Photo
White Pincushion Chaenactis artemisiifolia BFS, CP, Jep native Photo
California Thistle, Cobwebby Thistle Cirsium occidentale BFS, CP, Jep native Photo Several varieties may be present
Bull Thistle Cirsium vulgare BFS, CP, Jep non-native RSA882631
A small patch east of the lake was eliminated. A few additional plants that appeared south of the lake in the area burned in the 2018 Foothill Fire have been removed, but other may appear from time to time.
California Aster Corethrogyne filaginifolia BFS, CP, Jep native RSA611975 Formerly Lessingia filaginifolia. L. filaginifolia var. filaginifolia, L. filaginifolia var. bernardina, and L. filaginifolia var. peirsonii were all previously reported, but The Jepson Manual no longer recognizes any infraspecific taxa.
Australian Cotula, Southern Brass Buttons Cotula australis CP, Jep non-native RSA616147
San Diego Tarweed, Paniculate Tarplant Deinandra paniculata BFS, CP, Jep native RSA0171694 Listed as having “Limited distribution” in CNDDB. A single patch of approximately 100 plants spotted west of pHake Lake after the 2018 Foothill Fire has persisted.
African Daisy, Namaqualand Daisy Dimorphotheca sinuata BFS, CP, Jep non-native Photo Several plants were found in the area burned in the Foothill Fire and removed.
California Brittlebush Encelia californica BFS, CP, Jep native* Photo Had been planted behind the formed field house and has naturalized among the Redford Conservancy plantings.
Brittlebush, Incienso Encelia farinosa BFS, CP, Jep native Photo
Pine-Bush Ericameria pinifolia BFS, CP, Jep native RSA764413
Horseweed Erigeron canadensis BFS, CP, Jep native Photo Formerly Conyza canadensis.
Leafy Fleabane Erigeron foliosus var. foliosus BFS, CP, Jep native RSA869433
Golden-Yarrow Eriophyllum confertiflorum BFS, CP, Jep native RSA612655
Saw-toothed Goldenbush Hazardia squarrosa var. grindelioides BFS, CP, Jep native Photo The one known plant was bulldozed during the renovations for the Robert Redford Conservency, although others may be present.
Common Sunflower Helianthus annuus BFS, CP, Jep native Photo Uncommon. Has appeared from time to time.
Bristly Ox-Tongue Helminthotheca echioides BFS, CP, Jep non-native RSA817064 A single individual that appeared in the area burned by the Foothill Fire was removed.
Telegraph Weed Heterotheca grandiflora BFS, CP, Jep native RSA764412
Smooth Cat’s-Ear Hypochaeris glabra BFS, CP, Jep non-native RSA615307
San Diego Marsh-Elder, San Diego Povertyweed Iva hayesiana BFS, CP, Jep native RSA881791 Listed as “Imperiled” in CNDDB. Several plants are present on the banks of the wash.
Prickly Lettuce Lactuca serriola BFS, CP, Jep non-native Photo
California Goldfields Lasthenia californica subsp. californica BFS, CP, Jep native RSA616152 Has appeared transiently as a waif; not always present.
Hawkweed Layia Layia hieracioides BFS, CP, Jep native Photo, SD A small population near the north shore of pHake Lake first spotted in 2008 has since expanded. This represents a range extension for this species.
Scale-Broom Lepidospartum squamatum BFS, CP, Jep native RSA611981
Valley Lessingia Lessingia glandulifera var. glandulifera BFS, CP, Jep native RSA611968
Daggerleaf Cottonrose Logfia gallica CP, Jep non-native RSA616158
(as Filago gallica)
Formerly Filago gallica.
Stinknet Oncosiphon pilulifer BFS, CP, Jep non-native Photo Only a single plant (which was removed) has been detected to date.
Cudweed, Two-tone Everlasting Pseudognaphalium biolettii BFS, CP, Jep native UCR108175 Formerly Gnaphalium bicolor.
California Cudweed Pseudognaphalium californicum BFS, CP, Jep native RSA751109
(as Gnaphalium californicum)
Formerly Gnaphalium californicum.
Woolly-Heads, Slender Woolly-Marbles Psilocarphus tenellus CP, Jep native EN, R
Threadleaf Ragwort Senecio flaccidus var. douglasii BFS, CP, Jep native RSA444578
Common Groundsel Senecio vulgaris BFS, CP, Jep non-native RSA794414
Milk Thistle Silybum marianum BFS, CP, Jep non-native Photo In 2010 a large clump of Milk Thistle was removed from the in parkway on Mills Ave. It has not since recurred.
Prickly Sow Thistle Sonchus asper BFS, CP, Jep non-native Photo
Common Sow Thistle Sonchus oleraceus CP, Jep non-native LG
Wand Wirelettuce Stephanomeria virgata ssp. pleurocarpa BFS, CP, Jep native RSA611972
Rod Wirelettuce Stephanomeria virgata ssp. virgata BFS, CP, Jep native RSA823055
Everlasting Nest Straw Stylocline gnaphaloides BFS, CP, Jep native RSA612073
Cotton-Thorn Tetradymia comosa BFS, CP, Jep native RSA549566

Barberry Family (Berberidaceae)

Nevin’s Barberry Berberis nevinii BFS, CP, Jep native RSA792777

Birch or Alder Family (Betulaceae)

White Alder Alnus rhombifolia BFS, CP, Jep nativeL RSA611613

Borage Family (Boraginaceae)

Common Fiddleneck Amsinckia intermedia BFS, CP, Jep native POM612662
Formerly Amsinckia menziesii var. intermedia
Bearded Cryptantha Cryptantha barbigera CP, Jep native KH
Popcorn Flower Cryptantha intermedia BFS, CP, Jep native RSA612076
Redroot Cryptantha Cryptantha micrantha CP, Jep native KH
Minute-flowered Cryptantha Cryptantha micromeres CP, Jep native KH
Common Cryptantha Cryptantha muricata CP, Jep native RSA, JG
Prickly-Nut Cryptantha Cryptantha muricata var. denticulata CP, Jep native KH
Rigid Cryptantha Cryptantha nevadensis var. rigida CP, Jep native KH
Whispering Bells Emmenanthe penduliflora var. penduliflora BFS, CP, Jep native Photo
Hairy Yerba Santa Eriodictyon trichocalyx var. trichocalyx BFS, CP, Jep native RSA611616
Formerly placed in Hydrophyllaceae
Common Eucrypta Eucrypta chrysanthemifolia var. chrysanthemifolia BFS, CP, Jep native RSA612172
Formerly placed in Hydrophyllaceae.
Alkali Heliotrope, Seaside Heliotrope Heliotropium curassavicum var. oculatum BFS, CP, Jep native Photo Formerly Heliotropium curassavicum.
Comb-Bur, Narrow-toothed Pectocarya Pectocarya linearis ssp. ferocula CP, Jep native RSA616160B
Sleeping Combseed, Northern Pectocarya Pectocarya penicillata CP, Jep native RSA612072
Desert Bluebells Phacelia campanularia BFS, CP, Jep native* Photo In Mills Ave parkway. Not native to the BFS; probably planted in a wildflower mix.
Caterpillar Phacelia Phacelia cicutaria var. hispida BFS, CP, Jep native RSA869430 Formerly placed in Hydrophyllaceae.
Distant Phacelia Phacelia distans BFS, CP, Jep native RSA611615
Formerly placed in Hydrophyllaceae.
California Bells, Wild Canterbury Bells Phacelia minor BFS, CP, Jep native RSA612070
Formerly placed in Hydrophyllaceae.
Branching Phacelia Phacelia ramosissima BFS, Jep native RSA751108 Formerly in placed Hydrophyllaceae.
Lacy Scorpionweed, Tansy Phacelia Phacelia tanacetifolia BFS, CP, Jep native Photo, EN Formerly placed in Hydrophyllaceae.
Popcorn Flower Plagiobothrys canescens CP, Jep native RSA792783
Rough-stemmed Popcorn Flower Plagiobothrys collinus var. fulvescens CP, Jep native RSA616155

Mustard Family (Brassicaceae)

Common Sandweed Athysanus pusillus CP, Jep native RSA792772
Indian Mustard Brassica fruticulosa Jep non-native RSA612657
Reported in 1996 as a new introduction to North America, B. fruticulosa is now well established at the south foot of the San Gabriel and San Bernadino Mountains of Riverside, San Bernardino, and western Los Angeles Counties. A dense local infestation in the southwest corner of the ‘Neck’ was found in 1996. It has since spread to other parts of the BFS.
Black Mustard Brassica nigra BFS, CP, Jep non-native RSA611969 Uncommon at the BFS.
Turnip, Field Mustard Brassica rapa BFS, CP, Jep non-native RSA612660
Sahara Mustard Brassica tournefortii BFS, CP, Jep non-native RSA781424
First detected in 2012 in the parkway along Foothill Blvd., B. tournefortii subsequently spread inside the entry gate, by the Abilene Way gate, north of the infirmary, and inside the Foothill and Mills fences. As of 2021, it appears to have been eliminated inside the field station, but continues to appear in the parkway.
Bitter-cress, Toothwort Cardamine oligosperma BFS, CP, Jep native UC1595296
Shepherd’s Purse Capsella bursa-pastoris BFS, CP, Jep non-native Photo
Western Tansy-Mustard Descurainia pinnata BFS, CP, Jep native RSA792779
Garden Rocket Eruca vesicaria ssp. sativa BFS, CP, Jep non-native RSA831432
Short-pod Mustard Hirschfeldia incana BFS, CP, Jep non-native UCR108416
(both as Brassica geniculata)
Lesser Swine Cress Lepidium didymum BFS, CP, Jep non-native Photo Formerly Coronopus didymus.
Hairypod Pepperweed Lepidium lasiocarpum ssp. lasiocarpum CP, Jep native RSA799924
Robinson’s Peppergrass Lepidium virginicum var. robinsonii BFS, Jep native Photo, JS Listed as “Vulnerable” in CNDDB. The Jepson eFlora lumps L. virginicum var. robinsonii together with L. virginicum var. medium and L. virginicum var. pubesecens into L. virginicum var. menziesii.
Sweet Alyssum Lobularia maritima BFS, CP, Jep non-native RSA549565
Hedge Mustard Sisymbrium officinale BFS, CP, Jep non-native Photo
Indian Hedgemustard, Eastern Rocket Sisymbrium orientale CP, Jep non-native RSA748150

Cactus Family (Cactaceae)

Night-blooming Cereus, Peruvian Apple Cactus Cereus peruvianus BFS non-native Photo One planted outside the BFS Field House and a small group of plants naturalized from an adjacent yard on the eastern side of the ‘Neck’ have all been removed.
Cane or Snake Cholla Cylindropuntia californica BFS, CP, Jep native RSA612661
(as Opuntia parryi)
Formerly Opuntia parryi. The large genus Opuntia was subdivided into five genera; true chollas are in the genus Cylindropuntia, and O. parryi was renamed C. californica.
Indian-Fig, Mission Prickly-Pear Opuntia ficus-indica BFS, CP, Jep non-native Photo, JG
Prickly-Pear Opuntia littoralis BFS, CP, Jep native RSA748071

Pink Family (Caryophyllaceae)

Spreading Pygmyleaf Loeflingia squarrosa CP, Jep native RSA612084 Formerly Loeflingia squarrosa var. squarrosa. The Jepson Manual no longer recognizes any infraspecific taxa.
Four-leaved Allseed Polycarpon tetraphyllum CP, Jep non-native RSA616146
Sleepy Catchfly Silene antirrhina BFS, CP, Jep native Photo
Common Chickweed Stellaria media CP, Jep non-native EN

Goosefoot Family (Chenopodiaceae)

Four-wing Saltbush Atriplex canescens var. canescens BFS, CP, Jep native* Photo, SD A single large specimen near the southwest corner of the ‘Neck’, suspected to have naturalized to the BFS from the California Botanic Garden. Formerly in Amaranthaceae.
Pitseed Goosefoot Chenopodium berlandieri BFS, CP, Jep native UCD151063
Formerly in Amaranthaceae.
California Goosefoot Chenopodium californicum BFS, CP, Jep native RSA799915 Formerly in Amaranthaceae.
Russian Thistle Salsola australis BFS, CP, Jep non-native RSA831430
Russian Thistle Salsola tragus BFS, CP, Jep non-native RSA831431

Spiderflower Family (Cleomaceae)

Bladderpod Peritoma arborea BFS, CP, Jep native* RSA549570
(as Cleome isomeris)
(as Isomeris arborea)
Suspected to have naturalized to the BFS from the California Botanic Garden. Formerly Cleome isomeris and Isomeris arborea. Formerly placed in Capparaceae and Brassicaceae.

Morning-Glory Family (Convolvulaceae)

Island False Bindweed, Morning-Glory Calystegia macrostegia subsp. intermedia BFS, CP, Jep native RSA831434
Chaparral Dodder Cuscuta californica BFS, CP, Jep native RSA751110 Formerly placed in Cuscutaceae.

Stone Crop Family (Crassulaceae)

Pygmy-Weed Crassula connata CP, Jep native R
Moss Pygmyweed Crassula tillaea CP, Jep non-native RSA611986
Lance-leaved Dudleya Dudleya lanceolata BFS, CP, Jep native Photo

Cucumber Family (Cucurbitaceae)

Calabazilla, Buffalo Gourd Cucurbita foetidissima BFS, CP, Jep native Photo
Man-Root, Wild Cucumber Marah macrocarpa BFS, CP, Jep native RSA611611 Formerly Marah macrocarpus var. macrocarpus. The Jepson Manual no longer recognizes any infraspecific taxa.

Spurge Family (Euphorbiaceae)

California Croton Croton californicus BFS, CP, Jep native RSA611967
Dove Weed, Turkey Mullein Croton setiger BFS, CP, Jep native RSA611970
(both as Eremocarpus setigerus)
Formerly Eremocarpus setiger.
Rattlesnake Sandmat Euphorbia albomarginata CP, Jep native R Formerly Chamaesyce albomarginata.
Petty Spurge Euphorbia peplus BFS, CP, Jep non-native RSA612078
Thyme-leafed Spurge Euphorbia serpillifolia BFS, CP, Jep native Photo Formerly Chamaesyce serpyllifolia.
Castor Bean Ricinus communis BFS, CP, Jep non-native UCR107781

Legume (Bean & Pea) Family (Fabaceae)

Spanish Clover Acmispon americanus var. americanus BFS, CP, Jep native Photo Formerly Lotus unifoliolatus var. unifoliolatus.
Short-winged Deerweed Acmispon glaber var. brevialatus CP, Jep native UCR107830
(as Lotus scoparius var. brevialatus)
Formerly Lotus scoparius var. brevialatus.
Deerweed Acmispon glaber var. glaber BFS, CP, Jep native Photo Formerly Lotus scoparius var. scoparius.
San Diego Bird’s-foot-trefoil Acmispon micranthus CP, Jep native RSA612083
(as Lotus hamatus)
Formerly Lotus hamatus.
Bishop Lotus Acmispon strigosus BFS, CP, Jep native RSA612081
(as Lotus strigosus)
Formerly Lotus strigosus.
Bird-of-Paradise Shrub Caesalpinia gilliesii BFS, CP, Jep non-native RSA764417 Spreading into the east side of the ‘Neck’ from a neigbor’s yard.
Western Redbud Cercis occidentalis BFS, CP, Jep native* RSA766854 A single plant naturalized on the south shore of pHake Lake
Miniature Lupine Lupinus bicolor BFS, CP, Jep native RSA612177
Sky Lupine Lupinus nanus BFS, CP, Jep native* Photo In Foothill Blvd parkway. Not native to the BFS; probably planted in a wildflower mix.
Coulter’s Lupine Lupinus sparsiflorus BFS, CP, Jep native RSA817063
Arroyo Lupine Lupinus succulentus BFS, CP, Jep native Photo
Collar Lupine Lupinus truncatus CP, Jep native RSA612071
Sourclover Melilotus indicus BFS, CP, Jep non-native RSA615305
Mexican Palo Verde Parkinsonia aculeata BFS, CP, Jep non-native Photo, SD Naturalized in So. Cal.; one near the front gate and one in the wash near Lowell gate have been removed, but new plants keep appearing.
Honey Mesquite Prosopis glandulosa var. torreyana BFS, CP, Jep native* RSA748784
Truncate Sack Clover Trifolium depauperatum var. truncatum BFS, CP, Jep native RSA831435
Pinpoint Clover Trifolium gracilentum CP, Jep native RSA612065
Small-head Clover Trifolium microcephalum CP, Jep native RSA612082
Sweet Acacia Vachellia farnesiana CP, Jep non-native Photo Formerly Acacia farnesiana.

Beech Family (Fagaceae)

Oak (unspecified) Quercus sp. CP, Jep native* RSA792774 In experimental plots in East Field established by the Gamon lab
Coast Live Oak Quercus agrifolia var. agrifolia BFS, CP, Jep native RSA612079
Scrub Oak Quercus berberidifolia BFS, CP, Jep native RSA792836
Engelman Oak, Mesa Oak Quercus engelmannii CP, Jep native* SD Planted near the old infirmary but removed during the renovation of the Redford Conservancy.
Holly Oak Quercus ilex sp. CP, Jep non-native RSA792775 In experimental plots in East Field established by the Gamon lab
Valley Oak, Roble Quercus lobata BFS, CP, Jep native* Photo, SD Two deciduous oaks planted east of the entrance road just south of the infirmary.

Gentian Family (Gentianaceae)

Canchalagua, California Centaury, Charming Centaury Zeltnera venusta BFS, CP, Jep native UCR146987
(as Centaurium venustum)
Formerly Centaurium venustum

Geranium Family (Geraniaceae)

Filaree, Storksbill Erodium botrys CP, Jep non-native RSA751116
Red-stem Filaree Erodium cicutarium BFS, CP, Jep non-native RSA612653
White-stem Filaree, Greenstem Filaree Erodium moschatum BFS, CP, Jep non-native Photo

Gooseberry or Currant Family (Grossulariaceae)

Golden Currant Ribes aureum var. aureum CP, Jep native* RSA549572
Probably adventive from California Botanic Garden via birds as this variety is not native in the area
Golden Currant Ribes aureum var. gracillimum BFS, CP, Jep native RSA748783
White-Flowering Currant Ribes indecorum BFS, CP, Jep native RSA549567
Fuchsia-flowered Gooseberry Ribes speciosum BFS, CP, Jep native RSA549573
Not certain this is indigenous; it may have naturalized to the BFS from the California Botanic Garden.

Walnut Family (Juglandaceae)

Southern California Black Walnut Juglans californica BFS, CP, Jep native RSA764418
Formerly listed as Juglans californica var. californica, but the Jepson Manual no longer recognizes any infraspecific taxa.

Mint Family (Lamiaceae)

Henbit, Giraffe Head, Dead Nettle Lamium amplexicaule BFS, CP, Jep non-native RSA792771
Horehound Marrubium vulgare BFS, CP, Jep non-native RSA613095
White Sage Salvia apiana BFS, CP, Jep native RSA613094
Chia Salvia columbariae BFS, CP, Jep native Photo
Black Sage Salvia mellifera BFS, CP, Jep native Photo

Laurel Family (Lauraceae)

California Bay, California Laurel Umbellularia californica BFS, CP, Jep native* RSA613093 A group planted in front of the old infirmary has been removed, but some have naturalized on the site – likely from either that planting or California Botanic Garden.

Loosestrife Family (Lythraceae)

Hyssop Loosestrife Lythrum hyssopifolia CP, Jep non-native RSA616149

Mallow Family (Malvaceae)

Flannelbush Fremontodendron californicum BFS, CP, Jep native* UCR-108170 Formerly Fremontodendron californicum ssp. californicum; The Jepson Manual no longer recognizes any infraspecific taxa of this species. Formerly placed in Sterculiaceae.
Chaparral Mallow Malacothamnus fasciculatus BFS, CP, Jep native Photo, SD

Mahogany Family (Meliaceae)

Chinaberry Melia azedarach BFS, CP, Jep non-native Photo

Miner’s Lettuce Family (Montiaceae)

Red Maids Calandrinia menziesii BFS, CP, Jep native RSA799927
(as Calandrinia ciliata)
Formerly Calandrinia ciliata. Formerly placed in Portulacaceae.
Streambank Springbeauty Claytonia parviflora ssp. parviflora BFS, CP, Jep native RSA748782 Formerly placed in Portulacaceae.

Myrsine Family (Myrsinaceae)

Scarlet Pimpernel Lysimachia arvensis BFS, CP, Jep non-native RSA616150
(as Anagallis arvensis)
Formerly Anagallis arvensis. Formerly placed in Primulaceae.

Myrtle Family (Myrtaceae)

Blue Gum Eucalyptus globulus BFS, CP, Jep non-native Photo, SD
Red Iron Bark Eucalyptus sideroxylon BFS, CP, Jep non-native RSA748149

Four O’clock Family (Nyctaginaceae)

California Four O’Clock, Wishbone Bush Mirabilis laevis var. crassifolia BFS, CP, Jep native UCR108169
(as Mirabilis californica)
Formerly Mirabilis californica.

Olive Family (Oleaceae)

Desert Olive Forestiera pubescens BFS, CP, Jep native RSA778384
A single male plant grows on the north shore of pHake Lake.
Single-Leaf Ash or California Ash Fraxinus anomala or Fraxinus dipetala BFS
F. anomala:
CP, Jep
F. dipetala:
CP, Jep
native* Photo, EN Three trees near the College Ave gate were likely plants. Probably only one species of ash is present, but the species needs to be confirmed.
European Olive Olea europaea BFS, CP, Jep non-native RSA748148

Evening Primrose Family (Onagraceae)

California Sun Cup Camissoniopsis bistorta BFS, CP, Jep native RSA611985
(all as Camissonia bistorta)
Formerly Camissonia bistorta
Minature Sun Cup, Spencer Primrose Camissoniopsis micrantha BFS, CP, Jep native RSA817062 Formerly Camissonia micrantha
Four Spot Clarkia purpurea ssp. quadrivulnera BFS, CP, Jep native Photo
Elegant Clarkia Clarkia unguiculata BFS, CP, Jep native Photo
Willowherb Epilobium ciliatum CP, Jep native RSA766856
Water Primrose Ludwigia repens BFS, Jep nativeL RSA764407

Peony Family (Paeoniaceae)

California peony Paeonia californica BFS, CP, Jep native RSA792776

Poppy Family (Papaveraceae)

California Poppy Eschscholzia californica BFS, CP, Jep native Photo
Coulter’s Matilija Poppy Romneya coulteri BFS, CP, Jep native* RSA751106 Planted by the old infirmary; has subsequently naturalized.

Lopseed Family (Phrymaceae)

Southern Bush Monkey Flower Diplacus longiflorus BFS, CP, Jep native RSA612183
(as Mimulus aurantiacus)
Formerly included in Mimulus aurantiacus, but now considered a distinct species. Formerly placed in Scrophulariaceae.

Plantain Family (Plantaginaceae)

Blue Toadflax Nuttallanthus texanus BFS, CP, Jep native RSA616159
(as Linaria canadensis)
Formerly Linaria canadensis. Formerly placed in Scrophulariaceae.
Royal Penstemon Penstemon spectabilis BFS, CP, Jep native Formerly placed in Scrophulariaceae.
Royal Penstemon Penstemon spectabilis var. spectabilis CP, Jep native RSA612181
Formerly placed in Scrophulariaceae.

Sycamore Family (Platanaceae)

Western Sycamore Platanus racemosa BFS, CP, Jep native RSA613088

Phlox Family (Polemoniaceae)

Sapphire Woollystar Eriastrum sapphirinum BFS, CP, Jep native RSA751113
Sapphire Woollystar Eriastrum sapphirinum subsp. dasyanthum CP, Jep native RSA611971
Chaparral Gilia Gilia angelensis BFS, CP, Jep native RSA781423
Hollyleaf Navarretia Navarretia atractyloides BFS, CP, Jep native RSA751104

Buckwheat Family (Polygonaceae)

Leafy California Buckwheat Eriogonum fasciculatum var. foliolosum BFS, CP, Jep native RSA611976
Slender Wooly Wild Buckwheat Eriogonum gracile var. gracile BFS, CP, Jep native RSA611973
Leather Spineflower Lastarriaea coriacea CP, Jep native RSA612075
False Waterpepper Persicaria hydropiperoides CP, Jep nativeL RSA764408
(as Polygonum hydropiperoides)
Formerly Polygonum hydropiperoides
Dotted Smartweed Persicaria punctata BFS, CP, Jep nativeL RSA611980
(as Polygonum punctatum)
Formerly Polygonum punctatum
Woodland Threadstem Pterostegia drymarioides BFS, CP, Jep native RSA616157
Wild-Rhubarb, Canaigre Rumex hymenosepalus BFS, CP, Jep native RSA614148

Purslane Family (Portulacaceae)

Purslane Portulaca oleracea BFS, CP, Jep non-native Photo Has occurred as a waif.

Buttercup Family (Ranunculaceae)

Chaparral Clematis, Pipstem Clematis Clematis lasiantha BFS, CP, Jep native RSA792835
Cardinal or Scarlet Larkspur Delphinium cardinale BFS, CP, Jep native RSA612066
Parry’s Larkspur Delphinium parryi ssp. parryi BFS, CP, Jep native RSA612068
(no common name) Delphinium X inflexum BFS native Photo A fertile hybrid between Delphinium cardinale and Delphinium parryi ssp. parryi. A patch of these exists south of the lake. The name D. inflexum has been proposed but not accepted. Jepson eFlora mentions this hybrid under the listings for both Delphinium cardinale and Delphinium parryi.

Coffeeberry Family (Rhamnaceae)

Hoaryleaf Ceanothus Ceanothus crassifolius CP, Jep native* RSA792773 In experimental plots in East Field established by the Gamon lab
California Coffee Berry Frangula californica ssp. californica BFS, CP, Jep native RSA614147
(all as Rhamnus californica)
Formerly Rhamnus californica ssp. californica.
Spiny Redberry Rhamnus crocea BFS, CP, Jep native RSA549576

Rose Family (Rosaceae)

Birch-leaf Mountain-Mahogany Cercocarpus betuloides BFS, CP, Jep native Photo, R
Toyon, Christmas Berry Heteromeles arbutifolia BFS, CP, Jep native RSA611617
Mesa Horkelia, Wedge-Leaf Honeydew Horkelia cuneata var. puberula BFS, CP, Jep native RSA611622
NatureServe State Rank of S1 (Critically imperiled); listed as seriously endangered in California by the California Native Plant Society
Holly-leafed Cherry, Islay Prunus ilicifolia ssp. ilicifolia BFS, CP, Jep native RSA612184
P. ilicifolia ssp. ilicifolia hybridizes freely with P. ilicifolia ssp. lyonii, and intergrading forms may exist at the BFS.
Catalina Cherry Prunus ilicifolia ssp. lyonii BFS, CP, Jep native* Photo P. ilicifolia ssp. ilicifolia hybridizes freely with P. ilicifolia ssp. lyonii, and intergrading forms may exist at the BFS.
Small-leaved Rose Rosa minutifolia CP, Jep native* SD Planted near the old infirmary but removed during the renovation of the Redford Conservancy. On the California Endangered Species list, but protection does not apply to horticultural plantings.

Madder Family (Rubiaceae)

Goose Grass Galium aparine BFS, CP, Jep native RSA612077
San Diego Bedstraw Galium nuttallii subsp. nuttallii Jep native RSA799917

Willow Family (Salicaceae)

Fremont Cottonwood, Alamo Cottonwood Populus fremontii ssp. fremontii BFS, CP, Jep native* Photo, SD The only known individual near the Lowell gate was removed in the renovation of the Redford Conservancy
Gooding’s Black Willow Salix gooddingii BFS, CP, Jep nativeL RSA766853
Arroyo Willow Salix lasiolepis BFS, CP, Jep nativeL RSA612651

Soapberry Family (Sapindaceae)

Chinese Flame-Tree Koelreuteria bipinnata CP non-native RSA764420

Figwort Family (Scrophulariaceae)

Mediterranean Figwort, Nettle-leaved Figwort Scrophularia peregrina BFS, CP, Jep non-native RSA792782
Previously unreported in North America, S. peregrina had become an established weed throughout the Rancho Santa Ana Botanic Garden by 1996 (Ross and Boyd, Madroño 43(4): 436, July-Sept 1996) and has since spread to the BFS and other areas in Claremont. Native to Portugal and the Mediterranean region.
Wand Mullein Verbascum virgatum BFS, CP, Jep non-native RSA751105

Simarouba Family (Simaroubaceae)

Tree of Heaven Ailanthus altissima BFS, CP, Jep non-native RSA764419 All known individuals have been removed.

Nightshade Family (Solanaceae)

Jimson Weed Datura wrightii BFS, CP, Jep native Photo, JG
Manyflower Tobacco Nicotiana acuminata var. multiflora BFS, CP, Jep non-native Photo, SD
Tree Tobacco Nicotiana glauca BFS, CP, Jep non-native RSA611983
Indian Tobacco Nicotiana quadrivalvis BFS, CP, Jep native RSA748151
Douglas’ Nightshade Solanum douglasii BFS, CP, Jep native RSA613099
Purple Nightshade, Bluewitch Solanum umbelliferum BFS, CP, Jep native UCR107761
(as Solanum xanti)
Formerly listed as Solanum xanti, but we follow the classification of Knapp (PhytoKeys. 2013. 22: 1–432), which consolidates S. xanti within S. umbelliferum.

Nettle Family (Urticaceae)

Black-hair Nettle Hesperocnide tenella CP, Jep native RSA549581
Pellitory Parietaria hespera var. hespera CP, Jep native RSA615306
Dwarf Nettle Urtica urens BFS, CP, Jep non-native RSA549582

Vervain Family (Verbenaceae)

Southwestern Mock Vervain Verbena gooddingii BFS, CP, Jep native* Photo, SD Has occurred as a waif, likely originated from California Botanic Garden

Mistletoe Family (Viscaceae)

Big Leaf Mistletoe Phoradendron leucarpum ssp. macrophyllum BFS, CP, Jep native Photo, SD Formerly Phoradendron macrophyllum and Phoradendron serotinum ssp. macrophyllum

Grape Family (Vitaceae)

Wild Grape Vitis sp. BFS, CP, Jep native Photo, SD Because the few Vitis plants on the BFS have not borne fruit, species identification is difficult. Plants may be V. californica, V. girdiana, or a hybrid.

Caltrop Family (Zygophyllaceae)

Puncture Vine Tribulus terrestris BFS, CP, Jep native Photo, SD First detected in 2008 as an infestation in the northeast corner of the East Field. Despite eradication attempts, plants have appeared in the East Field, near the entry gate, and in the Foothill Blvd and Mills Ave parkways.


Agave Family (Agavaceae)

American Agave, Century Plant Agave americana BFS, CP, Jep non-native Photo Two infestations – one the wash south of Lowell gate and one in the lower ‘Neck’ have been eliminated. Formerly placed in Liliaceae.
Chaparral Yucca, Our Lord’s Candle Hesperoyucca whipplei BFS, CP, Jep native Photo Formerly Yucca whipplei. Formerly placed in Liliaceae.
California Soap Plant Chlorogalum pomeridianum var. pomeridianum BFS, CP, Jep native Photo Formerly placed in Liliaceae.

Arum Family (Araceae)

Duckweed Lemna valdiviana BFS, CP, Jep native RSA613092 Formerly placed in Lemnaceae.
Tongueshape Mud-Midget Wolffiella lingulata Jep native RSA613092
(in specimen of Lemna valdiviana)

Palm Family (Arecaceae)

Mexican Fan Palm Washingtonia robusta CP, Jep non-native S

Asparagus Family (Asparagaceae)

Bridal Creeper Asparagus asparagoides BFS, CP, Jep non-native Photo, LG

Sedge Family (Cyperaceae)

Tall Flatsedge Cyperus eragrostis CP, Jep nativeL RSA764405
Purple spikerush Eleocharis atropurpurea CP, Jep nativeL SD
Spikerush Eleocharis palustris CP, Jep nativeL RSA612180 Formerly listed as Eleocharis macrostachya, which closely resembles this species, but voucher specimen was determined to be palustris. Observed only in vernal pools to west of pHake Lake. These pools no longer exist.
Low Bulrush Isolepis cernua CP, Jep nativeL RSA766855
Naked-stemmed Bulrush Schoenoplectus sp. CP, Jep nativeL SD Formerly listed as Scirpus sp., but California Scirpus are now placed in Amphiscirpus, Bolboschoenus, Isolepis, Schoenoplectus, or Trichophorum.

Iris Family (Iridaceae)

Western Blue-eyed Grass Sisyrinchium bellum BFS, CP, Jep native RSA612174 Small populations were reported directly southeast of pHake Lake shore and south of fire road and near the trail at southwest corner of pHake Lake. Both appear to have perished in the 2014-2017 drought.

Rush Family (Juncaceae)

Baltic Rush Juncus balticus var. ater CP, Jep nativeL EN Clumped perennial with twisted stems and congested inflorescences afound on the margin of pHake Lake are tentatively identified as this species. [EN]
Toad Rush Juncus bufonius CP, Jep nativeL JG
Sonoran Rush,
Baja Rush
Juncus effusus ssp. austrocalifornicus Jep nativeL RSA611979
Formerly Juncus effusus var. brunneus.
Mexican Rush Juncus mexicanus CP, Jep nativeL RSA748779 Diffuse spreading plants with twisted stems, small open inflorescences, and leaves resembling stems are referred to this species, found only on the north shore of pHake Lake, near the roadway. [EN]
Wrinkled Rush Juncus rugulosus CP, Jep nativeL RSA611978

Grass Family (Poaceae)

Slender Wild Oat Avena barbata CP, Jep non-native RSA614152
Wild Oat Avena fatua CP, Jep non-native M
Ripgut Grass Bromus diandrus BFS, CP, Jep non-native RSA614150
Soft Chess, Soft Brome Bromus hordeaceus CP, Jep non-native RSA64154
Red Brome Bromus rubens BFS, CP, Jep non-native RSA614153
Formerly Bromus madritensis ssp. rubens
Giant Wild-Rye Elymus condensatus BFS, CP, Jep native Photo Formerly Leymus condensatus.
Rattail Sixweeks Grass Festuca myuros CP, Jep non-native RSA612080
(all above as Vulpia myuros)
Formerly Vulpia myuros var. hirsuta; varieties are no longer recognized
Hare Barley Hordeum murinum ssp. leporinum CP, Jep non-native R
Goldentop Lamarckia aurea CP, Jep non-native RSA707939
Crimson Fountaingrass Pennisetum setaceum CP, Jep non-native Photo, SD
Annual Bluegrass Poa annua CP, Jep non-native RSA616151
Annual Beard Grass, Rabbitfoot Grass Polypogon monspeliensis CP, Jep non-native RSA616145
Mediterranean Grass, Old Han Schismus Schismus barbatus CP, Jep non-native RSA616154

Pondweed Family (Potamogetonaceae)

Fennel-leaf Pondweed Stuckenia pectinata CP, Jep nativeL EN, JG Formerly Potamogeton pectinatus.

Brodiaea Family (Themidaceae)

Blue Dicks Dipterostemon capitatus ssp. capitatus BFS, CP, Jep native RSA612069
(All as Dichelostemma capitatum subsp. capitatum)
Formerly Dichelostemma capitatum subsp. capitatum. Formerly placed in Liliaceae and Asparagaceae.

Cattail Family (Typhaceae)

Broad-leaved Cattail Typha latifolia CP, Jep nativeL RSA611977


Pine Family (Pinaceae)

Knobcone Pine Pinus attenuata CP, Jep native* JG
Lacebark Pine Pinus bungeana BFS, CP non-native Photo
Coulter Pine Pinus coulteri CP, Jep native* EN, JG
Torrey Pine Pinus torreyana CP, Jep native* VS


Brake Family (Pteridaceae)

Goldback Fern Pentagramma triangularis BFS, CP, Jep native Photo, SD


native = California native, indigenous to the BFS
native* = California native planted or naturalized on the BFS; not part of the indigenous flora
non-native = Not native to California, according to Jepson eFlora
EN = Envicom, Field survey on the Robert J. Bernard Biological Field Station, September 28 and October 13, 1998
JG = Jeanne Marie Gural, Biological survey of the Robert J. Bernard Biological Field Station, Fall 1997 through Spring 2000
KH = Kristen Hasenstab, Rancho Santa Ana Botanic Garden
JS = Jane Strong, California Native Plant Society
LG = Leroy Gross, Rancho Santa Ana Botanic Garden
M = Field Station Manual, undated
R = Recon survey of the the Bernard Biological Field Station, Spring 1998
RSA = Rancho Santa Ana Botanic Garden plant list for the Robert J. Bernard Biological Field Station, Spring 1998
SD = Stephen Dreher, BFS Manager, 2001-2010
VS = Identification by Valerie Soza, 2001, or according to RSABG records (courtesy Valerie Soza, 2001)
Links to Photos and Descriptions:
BFS = Robert J. Bernard Biological Field Station
CP = CalPhotos, Digital Library Project, UC Berkeley. Clicking the CalPhotos link for a listing will return a display of all photos for that taxon. If you want to use any of these photos, be sure to check the guidelines for Using the Photos in CalPhotos
Jep = Jepson treatment and bioregional distribution from the Jepson eFlora, which parallels The Jepson Manual: Vascular Plants of California, 2nd Edition.
CDA = California Department of Food and Agriculture
POM = Pomona College Herbarium Collection, now combined with the Rancho Santa Ana Botanic Garden Herbarium
RSA = Rancho Santa Ana Botanic Garden Herbarium
SBBG = Santa Barbara Botanic Garden Herbarium
SDSU = San Diego State University Herbarium
UC = The University and Jepson Herbaria, University of California, Berkeley
UCR = University of California Riverside Herbarium